Megatron is a Decepticon, one of the lineal descendants of the military hardware robots created by the Quintessons on their factory world of Cybertron. Following a war between the Decepticons and the other robot race, the Autobots, the Decepticons were defeated by the Autobots' invention of transformation, which allowed them the advantange of stealth. The Autobot victory in the war began the Golden Age of Cybertron, but a viper lurked within this paradise, as the Decepticons too eventually developed transformation, eventually leading to the creation of Megatron. Gathering a small number of troops together, Megatron made a small strike on an outer city, killing the current Autobot leader. However, this event was not made public knowledge, and young robots like the naive Orion Pax still looked up to Megatron and his followers because of their new robot-mode flight powers. However, Pax soon learned the error of his ways when Megatron duped him into allowing him access to Energon warehouses, and Megatron fatally injured him in a scuffle. However, the ancient Autobot, Alpha Trion, reconstructed Pax into a battle hardy configuration - now, he was Optimus Prime, leader of the Autobots and Megatron's sworn opponent as the civil war erupted again.
Eventually, the war drained Cybertron of most of its energy, necessitating that both factions seek out new worlds and new sources of power. Megatron and his elite forces pursued the Autobots' craft, (referred to as the Ark in Beast Wars, following the ship's name from the G1 Marvel Comics) in their star cruiser (referred to as the Nemesis in Beast Wars) and attacked and boarded the craft, causing it to crash on a prehistoric planet, entombing all on the ship in emergency stasis as it crashed into a dormant volcano.
Four million years later, a volcanic eruption reactivated the Transformers and the war began again on the energy-rich world they found themselves on; Earth. After their first series of battles, the Autobots believed Megatron and the Decepticons destroyed when their new space cruiser crashed into the Pacific Ocean, but in reality they survived and reconstructed the cruiser to serve as a sub-oceanic staging base and bunker. Megatron re-established contact with Cybertron, and began construction of an intergalactic transport system called a "space bridge," although the early tests nearly 
resulted in his destruction when he was sucked into the portal and teleported to Cybertron. The technology was eventually stabilised, however, and used in a grand scheme to transport Cybertron through space into Earth's orbit, where the natural disasters its gravity wreaked created tremendous energy that Megatron and the Decepticons gathered. The Earth was saved, however, when Megatron's stockpiles of energy were detonated by the Autobots, the explosion forcing Cybertron out of orbit. Again, however, Megatron was able to cheat death, and was forced to team up with Optimus Prime in order to stop the machinations of his treacherous lieutenant, Starscream.
In his next scheme, Megatron absorbed all the separate powers of the other Decepticons into himself, and challenged Optimus Prime to one-on-one combat. With the added abilities of all his minions, Megatron easily defeated Prime in the battle, but when his deception was revealed, he and his followers were defeated in a group attack.Over the following years, Megatron's schemes to obtain Earth's energies continued. His assorted plans included various attempts to harness the power of Earth's core, thefts of several unstable energy devices from human scientists, the draining of energy from a time-lost prehistoric island,
and partnership with a human politician that allowed him to force the Autobots off of Earth, and actually allowed him to conquer Central City among many, many others.Eventually, however, Megatron turned his attention away from Earth, and succeeded in completely conquering Cybertron by the Earth year 2005. Learning the Autobots were preparing a strike against the planet, Megatron and the Decepticons intercepted the shuttle sent to Earth to acquire energy to power the raid, and unleashed a mighty assault on Autobot City.
In the course of the conflict, word was sent to Optimus Prime, who arrived the following morning after the slaughter and confronted Megatron in another one-on-one struggle. Megatron appeared beaten, but spying a discarded pistol, stalled for time by pleading for mercy as he attempted to reach it. The young Autobot Hot Rod then intervened, tackling Megatron, but the Decepticon was too powerful for him, seizing him and using him as shield as he grabbed the pistol and opened fire on Prime. The wounds were fatal and sent Prime to the grave - but not before he delivered his own final blow to Megatron, seriously damaging him and forcing the Decepticons to flee.On the return trip to Cybertron, it became necessary to jettison excess mass, or the Decepticons would be unable to reach the planet.
The wounded Decepticons were voted out, and set adrift in space, including Megatron. But this was not to be the end for the great slag-maker, as the world-devourer, Unicron, found him and recreated him... into Galvatron.
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